Stuart Little questions his ability after a grueling soccer match alongside George who kicked him with a soccer ball. He becomes even more downhearted after George's toy airplane gets broken in an accident because of him. On his way home from school, Stuart saves a female canary named Margalo who is being pursued by a peregrine falcon, and they become friends. But she is secretly working with Falcon to case and steal from households. When he presses her to find and take an object of value, or lose the sanctuary he promised her, she can't seem to concentrate on her assignment, as she is beginning to have tender feelings for Stuart. Falcon eventually loses patience and threatens to kill him if she doesn't deliver. Worried for his safety, she takes Eleanor Little diamond ring.
When the Littles see that the ring is missing, they think it has fallen down the sink. Stuart offers to be lowered down the drain on a string to get it, and nearly succeeds. When it breaks Margalo saves him, and his thanks to her only makes her feel even more guilty, so she decides to leave. When he can't find her, he assumes she has been kidnapped - and that Falcon is somehow involved. He leaves on a quest to rescue her with the household's reluctant cat Snowbell , but not before setting up a plan with George. Stuart and Snowbell decide to get information on Falcon's whereabouts, so they enlist the help of Monty. who tells them that Falcon's lair is at the disused observation deckof the nearby Pishkin Building. They hatch a plan to use balloons to get Stuart to the top, where he finds out that Margalo is Falcon's slave, and was forced to take the ring. He tries to save her, but Falcon captures him, and drops him from an immense height to his presumed death; however he lands on a soft trash bag in a garbage truck. Margalo and Snowbell are unaware of this. So when he goes to the top and asks her where Stuart is, she tells him that Falcon killed him.
On a garbage scow where he has ended up, Stuart blames himself for everything, and has almost lost all hope. But suddenly, he finds George's broken plane, fixes it up, and flies to save Margalo, who, having been freed by Snowbell, just fled from Falcon. The Littles, who have discovered his absence, (and that the plan Stuart made with George earlier was to lie to his parents) follow him by taxi as he begins an aerial adventure through the park, with Margalo at his side. Eventually, they lose Falcon, but he catches up and makes an attempt to kill Stuart, when he detaches the plane's upper wing, damaging the main one and causing it to enter a steep nose dive, which fails when Stuart recovers from the dive, nearly missing the Littles. He then realizes he can't run from Falcon, and lets Margalo off. He turns and flies the damaged plane in a kamikaze run while Falcon goes into an attack dive. He uses Mrs. Little's ring to temporarily blind him, and jumps out using a bandana as a parachute. The kamikaze attack works and Falcon is struck head on and defeated. Although he survives the attack, he falls out of the sky and lands in a garbage can that Monty is scavenging in, and is presumably eaten by him, but not before Stuart falls when his parachute is sliced apart by the propeller of the shattered plane, and then is rescued by Margalo. Stuart is congratulated by his family, and Margalo, who gives Mrs. Little her ring back, and Snowbell reunites with them as well. That evening, Margalo leaves with the other birds to migrate south, but not before saying goodbye to her friends. Stuart says the "silver lining" is that she'll be back in the spring, and his baby sister, Martha, says her first words: "Bye bye, birdie."
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